Borovice zhuštěná (Pinus densata) z tibetské části Číny je unikátním druhem mezi borovicemi světa: dosahuje nejvyšší výšky výskytu okolo 4 300 m n. m. (konkurované jediným druhem v tropech Ameriky), osídluje široké spektrum biotopů od subtropických údolních pralesů (subsp. tibetica) po skalnaté hřebeny velehor; jako druh s extrémními ekologickými adaptacemi a složitou evolucí se z hlediska molekulární genetiky stala nejstudovanější borovicí; její varieta pygmaea je nejmenší borovici světa (do 2 m výšky).

Použitá a citovaná literatura:

ARMITAGE, F. B., et al. Pinus kesiya Royle ex Gordon syn. P. khasya Royle; P. insularis Endlicher. Commenwealth Forestry Institute, University of Oxford, 1980.
BUSINSKÝ, Roman. A reassessment of Tibetan hard pine, Pinus densata, on the basis of morphology and biogeography of the P. kesiya complex (Pinaceae). Harvard papers in botany, 2008a, 13.1: 1-28.
BUSINSKÝ, Roman. The genus Pinus L., pines: contribution to knowledge. A monograph with cone drawings of all species of the world by Ludmila Businská. Acta Pruhoniciana, 2008b, 88: 1-126.
BUSINSKÝ, Roman; FRANTÍK, Tomáš; VÍT, Petr. Morphological evaluation of the Pinus kesiya complex (Pinaceae). Plant systematics and evolution, 2014, 300.2: 273-285.
CRITCHFIELD, William Burke; LITTLE, Elbert Luther. Geographic distribution of the pines of the world. US Department of Agriculture, Forest Service, 1966.
FARJON, Aljos. World checklist and bibliography of conifers. Royal Botanic Gardens, Kew, Richmond, UK, 2001.
FU, Liguo; LI, N.; MILL, R. R. Pinaceae. In: WU, Z. Y.; RAVEN P. H. (eds.), Flora of China 4 (Cycadaceae through Fagaceae). Science Press, Beijing & Missouri Bot. Gard. Press, St. Louis, 1999, 11-52.
GAUSSEN, H. Les Gymnospermes actuelles et fossiles. Fascicule VI, Chapitre XI. Généralités, Genre Pinus. Trav. Lab. Forest.Toulouse [Tome 2, Sect. 1], 1960, Vol. 1 [Part. 2]: 1–272.
GERNANDT, David S., et al. Phylogeny and classification of Pinus. Taxon, 2005, 54.1: 29-42.
GROTKOPP, Eva, et al. Evolution of genome size in pines (Pinus) and its life-history correlates: Supertree analyses. Evolution, 2004, 58.8: 1705-1729.
CHEN, Kangming, et al. Phylogeography of Pinus tabulaeformis Carr.(Pinaceae), a dominant species of coniferous forest in northern China. Molecular Ecology, 2008, 17.19: 4276-4288.
LAW, Y. W., et al. Pinus. In: CHENG, W. C.; FU, L. K. (eds.). Flora Reipublicae Popularis Sinicae, 1978, 7: 204-281.
LI, De-Zhu. A reassessment of Pinus subgen. Pinus in China. Edinburgh Journal of Botany, 1997, 54.3: 337-349.
LIU, Zhan-Lin; CHENG, Cheng; LI, Jianfang. High genetic differentiation in natural populations of Pinus henryi and Pinus tabuliformis as revealed by nuclear microsatellites. Biochemical systematics and ecology, 2012, 42: 1-9.
MA, Xiao-Fei; SZMIDT, Alfred E.; WANG, Xiao-Ru. Genetic structure and evolutionary history of a diploid hybrid pine Pinus densata inferred from the nucleotide variation at seven gene loci. Molecular Biology and Evolution, 2006, 23.4: 807-816.
MAO, Jian-Feng; WANG, Xiao-Ru. Distinct niche divergence characterizes the homoploid hybrid speciation of Pinus densata on the Tibetan Plateau. The American Naturalist, 2011, 177.4: 424-439.
MIROV, Nicholas Tiho, et al. The genus pinus. The genus Pinus. Ronald Press, 1967.
MYBURG, H.; HARRIS, S. A. Genetic variation across the natural distribution of the south east Asian pine, Pinus kesiya Royle ex Gordon (Pinaceae). Silvae Genetica (Germany), 1997, 46.5: 295-301.
PRICE, Robert A.; LISTON, Aaron; STRAUSS, Steven H. Phylogeny and systematics of Pinus. Ecology and biogeography of Pinus. In RICHARDSON, David M.; RUNDEL, Philip W. Ecology and biogeography of Pinus: an introduction. Ecology and biogeography of Pinus, 1998, 49-68.
WANG, Xiao-Ru; SZMIDT, Alfred E. Chloroplast DNA-based phylogeny of AsianPinus species (Pinaceae). Plant systematics and Evolution, 1993, 188.3-4: 197-211.
WANG, Xiao-Ru; SZMIDT, Alfred E. Hybridization and chloroplast DNA variation in a Pinus species complex from Asia. Evolution, 1994, 48.4: 1020-1031.
WANG, Xiao-Ru, et al. Phylogenetic relationships of Eurasian pines (Pinus, Pinaceae) based on chloroplast rbcL, matK, rpl20-rps18 spacer, and trnV intron sequences. American Journal of Botany, 1999, 86.12: 1742-1753.
WANG, Baosheng, et al. Colonization of the Tibetan Plateau by the homoploid hybrid pine Pinus densata. Molecular ecology, 2011, 20.18: 3796-3811.
WU, C. L. The taxonomic revision and phytogeographical study of Chinese pines. Acta Phytotaxonomica Sinica, 1956, 5: 131–164, Pl. 24–27.
YING, Junsheng, et al. Atlas of the Gymnosperms of China. China Science and Technology Press, 2004.

Pinus densata from the Tibetan part of China is a unique species among world pines – it reaches the highest elevation of approximately 4 300 m (rivalled by the only species in tropical America), and colonises the broad biotop spectrum from subtropical valley forests (P. d. subsp. tibetica) to the rocky ridges of high mountains; it is the pine that has been most researched by mo­lecular genetics as a species with extreme ecological adaptations and complex evolution. Pinus densata var. pygmaea is the smallest pine in the world (up to 2 m).